Taxonomy of Rudists from the campanian transgressive sediments of Brašljevica, Donje Orešje and Sv. Martin, northern Croatia

Rudistler, Hırvatistan’ın kuzeybatısında yer alan üç lokalitedeki transgresif Üst Kretase çökellerinde yaygın olarak bulunan fosillerdir. İlk ortaya çıkışları, transgresif istifin başlangıcında yer alan yüzertaş katmanlarına karşılık gelir. Rudist topluluğu içersinde, Vaccinites Fischer 1887, Hippurites Lamarck 1801 ve Hippuritella Douvillé 1908, genel veya cinse ait morfolojik parametreler ile birbirlerinden ayrılabilir. Donje Orešje ve Brašljevica lokalitelerinde çökellerin yaşı, rudist kavkılarının Sr-izotop içeriği ve rudistli tortulları üzerleyen sedimanlarda saptanan mikro ve nannofosil topluluklarından elde edilmiştir. Bu yaş verilerine göre transgresif tortulların yaşı Erken Kampaniyen’dir. Kavkıların enine kesitlerinde tanımlanan morfolojik parametreler (ligament, piliyeler, iç çap, dış kavkının iç konturunun uzunluğu, bu uzunluk ile piliyelerin süturları arasındaki mesafenin oranı) Hippuritidae, Gray 1848 ailesinde farklı türlerin tanımlanmasını olanaklı kılar. Çalışılan Vaccinites, Hippurites ve Hippuritella türlerinin enine kavkı kesitleri farklı ölçülmüş morfolojik parametre değerleri ve oranlar sunar. Bu değer ve oranlar, Vaccinites cornuvaccinum (Bronn 1831)’un tüm lokalitelerde, V. giganteus (d’Hombres) ve V. vesiculosus (Woodward 1855)’un Brašljevica ve Donje Orešje’de ve Hippurites vidaliMatheron 1880 ve Hippuritella lapeirousei (Goldfuss 1840)’nun Donje Orešje’de olası taksonomik tanımlamaları için bir zemin oluşturan Kümeleme analizi’ne (Ward ve Unweighted grup çifti metodu) tabi tutulmuştur.

Brašljevica, Donje Orešje ve Sv. Martin’de (Kuzey Hırvatistan) kampaniyen transgresif tortullarının Rudist taksonomisi

Rudists are common fossils in transgressive Upper Cretaceous deposits in three localities placed in northwestern part of Croatia. Their appearances coincide with the beginning of a transgressive sequence through floatstones-thickets. Within the rudist community, Vaccinites Fischer 1887, Hippurites Lamarck 1801 and Hippuritella Douvillé 1908 could be differentiated according to their general or generic morphological elements. The age of deposits was obtained from the Sr-isotope composition of the rudist shells as well as the micro- and nannofossil community from sediments overlying the rudists-bearing sediments at the Donje Orešje and Brašljevica localities. According to this data the age of the transgressive sediments are determined as Early Campanian. Morphological elements (ligamental ridge, pillars, inner diameter, length of the contour around the inner margin of the outer shell layer, and the ratio between this length and the distance between the sutures of the pillars) from the transverse shell sections permit possible definition of different species within the family Hippuritidae, Gray 1848. Transverse shell sections of analyzed Vaccinites, Hippurites and Hippuritella specimens show different values of measured morphological elements as well as their ratios which, subjected to cluster analysis (Ward's and Unweighted pair-group methods), represent grounds for possible taxonomic determination of the species Vaccinites cornuvaccinum (Bronn 1831) at all localities; V. giganteus (d'Hombres-Firmas 1838) and V. vesiculosus (Woodward 1855) at Brašljevica and Donje Orešje; as well as Hippurites vidali Matheron 1880 and Hippuritella lapeirousei (Goldfuss 1840) at Donje Orešje.

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