A conspectus of Scorzonera s.l. in Turkey

A comprehensive taxonomic study based on comparative morphology of Scorzonera, here maintained in its wide sense, is presented for the territory of Turkey. This study has produced several changes of classification at sectional and species ranks. An updated list of Scorzonera taxa occurring in Turkey, along with their infrageneric classification, is provided. A new section, S. sect. Anatolia Makbul & Coskunç., is described. Chromosome numbers, threat categories, and distribution maps are given for 6 endemic taxa (S. boissieri, S. karabelensis, S. longiana, S. sandrasica, S. ulrichii, and S. zorkunensis) placed in or transferred to the newly described section. A revised identification key to all Scorzonera species in Turkey is presented.

A conspectus of Scorzonera s.l. in Turkey

A comprehensive taxonomic study based on comparative morphology of Scorzonera, here maintained in its wide sense, is presented for the territory of Turkey. This study has produced several changes of classification at sectional and species ranks. An updated list of Scorzonera taxa occurring in Turkey, along with their infrageneric classification, is provided. A new section, S. sect. Anatolia Makbul & Coskunç., is described. Chromosome numbers, threat categories, and distribution maps are given for 6 endemic taxa (S. boissieri, S. karabelensis, S. longiana, S. sandrasica, S. ulrichii, and S. zorkunensis) placed in or transferred to the newly described section. A revised identification key to all Scorzonera species in Turkey is presented.

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  • Appendix. List of examined taxa. Scorzonera ahmet-duranii: C2 Muğla: Köyceğiz, Sandras mountain, Topuklu-fire tower, Pinus clearance, 1655 m, 23.06.2010, Makbul & Coşkunçelebi 230 (RUB, KTUB). Scorzonera bella: C10 Hakkâri: Cilo mountain in
  • Diz stream, 1730 m, 06.08.1954, Davis & Polunin 23925
  • (LE). Scorzonera boissieri: B6 Kayseri: Sarız, Yeşilkent, Binboğa mountains, Tekke rock, mountain steppe, 1879 m, 07.2010, Makbul & Coşkunçelebi 250 (RUB, KTUB);
  • Maraş: Göksun, Berit mountain, above Çavdar, rocky slopes, 2203 m, 14.07.2010, Makbul & Coşkunçelebi 247
  • (RUB, KTUB). B6 Kayseri: Sarız, Yalak, Binboğa mountains, Tekke rocky places, steppe, 2000–2200 m, 001992, Aytaç & Duman 5157 (HUB); Kayseri: Sarız,
  • Keklikoluk rock, Işık mountain, Binboğa mountain, mountain steppe, 2400 m, 12.09.1991, Aytaç 4451 (HUB);
  • Kayseri: Binboğa mountain, Körkuyu-Sıcak plateau, stony steppe, 2400–2600 m, 21.07.1992, Aytaç & Duman 5444
  • (GAZI); Maraş: Göksun, Ericek, Berit mountain, above Çavdar plateau, 2200–2250 m, 29.06.1992, Aytaç & Duman 5004 (GAZI). Scorzonera coriacea: B3 Isparta:
  • Şarkikaraağaç, Kızıldağ National Park, 1600–1700 m, 01994, B.Mutlu 579-928 (HUB); C3 Konya:
  • Derebucak, Çamlık, Kızıldağ, serpentine places, 1400 m, 02009, N 37 21 869 E 031 40 501, A. Duran 8349 & B.
  • Doğan (holo.: KNYA; iso.: GAZI, ANK, HUB, Selçuk Univ., Herbarium of the Faculty of Education); 1445 m, N 37 21 028-E 031 39 082, 30.05.2005, E.Hamzaoğlu 3707 &
  • Aksoy (KNYA); Suluin road, serpentine places, 1300 m, N 37 21 299 E 031 36 467, 11.06.2008, A. Duran 8042
  • (KNYA); Konya: Derebucak, Çamlık road, opposite the fountain, Pinus clearance, 1401 m, 12.06.2009, Makbul &
  • Coşkunçelebi 144 (RUB, KTUB); C3 Isparta: Şarkikaraağaç, Kızıldağ National Park, around the hill, subalpine, alpine, 1599 m, 14.07.2012, Makbul & Coşkunçelebi 354 (RUB, KTUB). Scorzonera davisii: C10 Hakkari: 30 km from
  • Hakkari to Başkale, stony slopes, 1430 m, 19.07.2011,
  • Makbul & Coşkunçelebi 320 (RUB, KTUB); Van: Başkale– Hakkari road, 20–25 km from Hakkari, Zap gorge, roadside, slopes, 1533 m, 19.07.2011, Makbul &
  • Coşkunçelebi 315 (RUB, KTUB). Scorzonera elata: A2 (A) İstanbul: Kartal, Cevizli, Dragos mountain, hill, Pinus forest, 28.04.1968, A. Çır. and O. Sut. (ISTF); İstanbul:
  • Büyük island, 0–100 m, Sibthorp; Bursa: Bursa surroundings, Sibthorp; A4 Kastamonu: Daday, Juniperus forest, 700 m, 26.06.1980, Ketenoğlu 792 (ANK); A5
  • Kastamonu: Tosya, Kavakçeşme, Sint. 1892: 4918b; Tokat:
  • Zile, Akçakeçeli village, Quercus forest, calcareous stones, 850–950 m, 24.07.1993, Dönmez 3789 (HUB); B1
  • Balıkesir: Ayvalık, Pınar island, 10 m, 10.05.1996, K.
  • Akpınar (ISTE); İzmir: İzmir surroundings, Montbret; İzmir: Bornova–Manisa 15 km, hill 430 m, 08.07.1993, Ö.
  • Seçmen 4291 (EGE); İzmir: Bayraklı, meadows, 200–300 m 001933, O. Schwarz (EGE); İzmir: Selçuk–Kuşadası,
  • Samsun mountain, 24.06.1965, Regel (EGE); Manisa: 20 km S of Manisa, Gediz, 500 m, 26.07.1962, M. & D. Zohary;
  • B3 Eskişehir: Türkmen mountain, Arban stream, Üçsaray, P. sylvestris clearance, 16.06.1976, T. Ekim, (ISTE); Konya:
  • Akşehir, above Atakent, 1080–1200 m, 28.05.1974, G.
  • Dökmeci and Y. Doğantaş (ISTE); B4 Ankara: Mamak, Kıbrıs village valley, Cehrelik, grassland, 1300–1350 m, 002005, Aslan 2419 (GAZI); Ankara: Ayaş Beli, Gazi
  • Mustafa Kemal Forest, forest clearance, 1100–1200 m, 001986, Aytaç 2179 (GAZI); B6 Sivas: 12 km N of Sivas, 1600 m, Sorger 60-18-17; B7 Sivas: Divriği, 1000 m, Bornm 1893: 242a; Erzincan: Salihli, 1250–1300 m, Hub.-Mor. 9117; C1 Aydın: Priene residues, 250 m, 25.05.1962, D. 34975 (E); Aydın: Samsun mountain, Regel (EGE 8296);
  • İzmir: Selçuk, 300 m, Sorger 63-13-4; Muğla: Bodrum, S of Turgut Reis, W foot of Doru mountain, 16.05.1984, E.
  • Tuzlacı (ISTE); C2 Aydın: from Aydın to Muğla 72 km, 330 m, Hub.-Mor. 13193; Antalya: Akdağ, 1400 m, Sorger 67- 22-75; Burdur: 3 km N of Dirmil, 1250–1300 m, Hub.
  • Mor. 9115; Denizli: Denizli–Kaçık Beli road, Karatepe,
  • Gökkaya plantation area, 750 m, 07.06.1973, A.Baytop and T.Tuzlacı (ISTE); Denizli: Babadağ, Antalya–Tavas junction, roadside, 500 m, 20.07.1997, S. Oluk 2656 (EGE);
  • Muğla: Küçük Kargı, from Fethiye to Köyceğiz, Bozakman & Fitz 344; Muğla: Ortaca, Tepearası, Eskiköy, Pinus brutia forest, 60–80 m, 25.05.1991, Vural 5873, (HUB); Muğla:
  • Köyceğiz, Kavakarası village, maquis, 20 m, 18.05.1991,
  • Güner 9047 (HUB, GAZI); Muğla: Köyceğiz, Hamitköy,
  • Domuzdireği hill, maquis, 10–30 m, 21.05.1991, Güner 9169 (HUB); Muğla: Köyceğiz, Ekincik village, Sandallı hill, 400–600 m, 16.06.1991, Güner 9394 (HUB); Muğla:
  • Köyceğiz, Beybası village, Süpürgelik hill, maquis, 150 m, 01991, Güner 8994, (HUB); Muğla: Dalaman, from
  • Dalaman to Köyceğiz 10 km before, P. brutia forest, roadside, 152 m, 25.05.2010, Makbul & Coşkunçelebi 204
  • (RUB, KTUB); C3 Antalya: Tahtalı mountain, Tekirova, 30 m, Hub.-Mor. 9983; Antalya: Adrasan, Çavuş village, Sazak road, 15 m, 15.05.2009, Eustace Ann 118 (E); Antalya:
  • Akseki–Güzelsu, Serebil area, Pinus brutia forest, 1100 m, 001996, Duran 4136, (GAZI); C4 Antalya: Alanya,
  • Çetik 124; Antalya: Alanya, Mahmutlar, Kuşkayası, Pinus nigra forest, 1371 m, 04.07.2010, Makbul & Coşkunçelebi 236 (RUB, KTUB); C6 Maraş: 3 km S of Süleymanlı, shrubs, 900 m, 16.06.1981, Yıldız 2811 (HUB ). Scorzonera hieraciifolia: B4 Konya: Gölyazı plateau, salty places, 912 m, 06.2009, Makbul & Coşkunçelebi 138 (RUB, KTUB);
  • B4 Aksaray: Tuz lake, Aksaray, Eskil-lake, swamp, Juncus community, 940 m, 02.06.1998, Aydoğdu (GAZI); Konya:
  • Cihanbeyli, 800 m, Hub.-Mor. 14690; B5 Kayseri: İncesu
  • Develi, 1200 m, salty swamp, 29.08.1957, D. 32766 (Typus)
  • (E); Kayseri: İncesu-Develi, 1200 m, D. 32766; Kayseri:
  • Sultan reeds, Yahyalı, Ovaçiftliği village, Kuş museum, protected area, 1071 m, 10.07.1993, Yıldırımlı 16171
  • (HUB); Kayseri: Yahyalı, Sultan reeds, Tuzla, 03.07.1982,
  • Sümbül 1475 (HUB); Kayseri: Yahyalı-Sultan reeds, Tuzla, 001982, Demirkuş 1913 (HUB); Niğde: Aksaray,
  • Birand & M.Zohary 2819; B6 Sivas: Tödürge lake, Zara
  • Hafik, 1350 m, Hub.-Mor. 13204; B6 Sivas: Tödürge lake, S of Zara-Hafik, salty places, 1301 m, 06.06.2009, Makbul &
  • Coşkunçelebi 127 (RUB, KTUB). Scorzonera inaequiscapa: A7 Giresun: 10–15 km from Alucra to Şiran, Hacıhasan village, grassy and rocky area, 1670 m, 13.06.2005, Makbul 79 (RUB, KTUB); Giresun: Alucra–Şiran, Hacıhasan village, grassland, sandy and rocky area, 1661 m, 02011, Makbul & Coşkunçelebi, 298 (RUB, KTUB);
  • A8 Bayburt: 2.5 km from Bayburt to İspir, roadside, 1506 m, 05.2005, Makbul 68 (RUB, KTUB); B5 Yozgat:
  • Akdağmadeni forest station, 06.06.1960, Curtis 177; B7
  • Tunceli: Ovacık, Karagöl valley, Munzur mountain, 1400 m, 005.1979, Yıldırımlı 1347 (HUB). Scorzonera karabelensis: C2 Muğla: Fethiye, Fethiye–Korkuteli road, clearance of Pinus forest, rocky chalky slopes, 1114 m, 02012, Makbul & Coşkunçelebi 346 (RUB, KTUB).
  • Scorzonera longiana: C4 Antalya: Gazipaşa, Çayıryakası plateau, 1700 m, 14.07.1983, Sümbül 2290 (HUB); Antalya:
  • Gazipaşa, from Çobanlar village, plateau to Oyuklu plateau, 1800–2000 m, 11.07.1983, Sümbül 2239 (types HUB, E). Scorzonera longiana: C3 Antalya: Gazipaşa,
  • Çayıryaka plateau, stony meadows, 1727 m, 25.07.2010,
  • Makbul & Coşkunçelebi 255 (RUB, KTUB). Scorzonera pacis: C6 Hatay: Antakya, Karaali, maquis, 410 m, 02011, S. Yıldız 551 & S. Kayıkçı (holo.: GAZI, iso.: HUB, MKU). Scorzonera phaeopappa: C3 Antalya:
  • Akseki, Yaylacık hill, 1900 m, Quezel et al.; C5 Adana:
  • Adana–Ceyhan, 6 km E of Ceyhan stream, Coode & Jones 3571; Adana: Mısır–Ceyhan, 25 m, 14.04.1956, D. 26069
  • (ANK); Adana: Mısır, 11.04.1934, Balls 717 (ANK); Adana: 8 km from Adana to Kozan, calcareous soils, 150 m, 01957, D. 26628 (E); Niğde: Ala mountain, Çukurbağ–
  • Narpız, 1900–2050 m, Parry 72; C6 Gaziantep: Dülük
  • Baba, 7 km N of Gaziantep, 1100 m, 12.05.1957, D. 27879
  • (ANK, E); Gaziantep: Dülük Baba, Gaziantep, 1250 m,
  • Haradj. 1217; Gaziantep: Dülük Baba, picnic area, Pinus forest, 1135 m, 15.05.2010, Makbul & Coşkunçelebi 174
  • (RUB, KTUB); Hatay: Kırıkhan, Antalya–İskenderun, 5 km from Kırıkhan junction, roadside, 385 m, 16.05.2010,
  • Makbul & Coşkunçelebi 177 (RUB, KTUB); Maraş: Ahır mountain, Yalnız Ardıç backs, 1300–1500 m, 22.05.1992,
  • Aytaç & Duman 4668 (HUB); Maraş: Engizek mountain,
  • Aksu, Juniperus excelsa clearance, 1000–1200 m, 01987, Duman 2919 (GAZI); Maraş: Ahır mountain,
  • Akdere, 1300–1600 m, 03.05.1991, Aytaç 3564 (GAZI); C7
  • Urfa: Ceylanpınar, Sorkah, 470–490 m, 14.04.1995,
  • Adıgüzel 1895 (GAZI); C8 Batman: Bismil, 5 km from steppe area to hill, 610 m, 25.04.2009, K 37 81 06-D 40 72 53, Aslan et al. 3210 (GAZI); Mardin: 24 km from
  • Diyarbakır to Mardin, 1000 m, D. 28735; Urfa: Ceylanpınar,
  • Hausskn. Scorzonera pisidica: C2 Burdur: Tefenni, Quercus coccifera, 3 km from Altınyayla, 1200–1270 m, 01948, Hub.-Mor. 8485 & Reese (holo Hb. Hub. Mor.);
  • Burdur: 6 km S of Dirmil, 1600–1650 m, Hub.-Mor. 8486
  • & Reese; Burdur: Gölhisar-Dirmil, 1270 m, 17.06.1981,
  • Nydegger 16244 (tip) (E); Burdur: Yeşilova, S of Salda lake,
  • P. nigra and Quercus clearance, 1170–1200 m, 11.07.1993,
  • Duman 5080 (GAZI); C2 Denizli: Honaz, roadside, stony field, Juniperus clearance, 1743 m, 13.07.2012, Makbul &
  • Coşkunçelebi 353 (RUB, KTUB); Muğla: Köyceğiz, Sandras mountain, climbing from Beyağaç to Sandras mountain, Yumaklı area, Pinus sp. forest, 1282 m, 02010, Makbul & Coşkunçelebi 229 (RUB, KTUB);
  • Muğla: Köyceğiz, Sandras mountain, Topuklu–Yangın fire tower, Pinus sp. forest, 1655 m, 23.06.2010, Makbul &
  • Coşkunçelebi 231 (RUB, KTUB); Muğla: Beyağaç–Sandras mountain, P. nigra forest, 1320 m, 16.06.2001, Varol 3895
  • (GAZI). Scorzonera pygmaea: A2 Bursa: Uludağ, Juniperus clearance, 2159 m, 24.07.2010, Makbul & Coşkunçelebi 271 (RUB, KTUB); A4 Kastamonu: Ilgaz mountain, around TV transmitter, alpine, 2055 m, 02.08.2009,
  • Makbul 165 (RUB, KTUB); A5 Kastamonu: Ilgaz mountain, back hills of the TV transmitter, steppe, 1998 m, 08.2011, Makbul & Coşkunçelebi 339 (RUB, KTUB).
  • A2(A) Bursa: Uludağ, 19.08.1850, Clementi; 2400 m,
  • Sorger 68-54-23; Bursa: Uludağ, Karagöl road, 24.08.1971,
  • A. & T. Baytop (ISTE); Bursa: Uludağ, alpine, 13.08.1953,
  • A. & T. Baytop (ISTE); 29.08.1950, A. & T. Baytop (ISTE);
  • A2 Bursa: Uludağ, 2380–2580 m, 13.09.1949, D. 14846- 14862a (E); Bursa: Uludağ, 29.07.1968, Pamukçuoğlu
  • (HUB); Bursa: Uludağ, NW of hill, calcareous stones, 2300–2500 m, 05.08.1993, Y. Gemici 8284 (EGE); Bursa:
  • Uludağ, around the hill, stream bed, 1950 m, 27.07.1984,
  • Ö. Seçmen 11 (EGE); Karabük: peak of Keltepe, 1950 m, 01984, Demirörs 1263 (ANK); Kastamonu: Ilgaz mountain, limy stones, 2200 m, 28.07.1962, D. 21577- 38376 (E); Kastamonu: Ilgaz mountain, Pinus forest, 2000 m, 06.1969, Darrah 28 (E); Kastamonu: Ilgaz mountain,
  • K. Hacet mountain, S slopes of alpine, limy, 2400 m, 01982, Akman et al. 12178 (ANK, EGE); A4
  • Zonguldak: Karabük-Keltepe, 1950 m, calcareous hills, 001962, D. 39814 (E); B2 Kütahya: Gediz, Şaphane mountain, calcareous stones, 1700–1900 m, 17.06.1993,
  • Y.Gemici 7695 (EGE); C3 Isparta: Dedegöl Mountain, 1600–2400 m, Sorger 70-46-24. Scorzonera rigida: B7
  • Erzincan: above Ahmediye, NW slopes, alpine, stony places, 2306 m, 11.07.2009, Makbul 156 (RUB, KTUB); C4
  • Konya: Ereğli, Kayasaray village, Düğünlük stream, calcareous stones, 1910 m, 05.07.2010, Makbul 240 (RUB, KTUB). B7 Erzincan: Sipikör mountain, 1990 m, Hub.
  • Mor. 13194; B9 Bitlis/Van: 10 km SE of Pelli, calcareous slopes, 2600 m, 08.07.1954, D. 22544 (ANK, E); Van:
  • Başkale, İspiriz mountain, stony slopes, 3200 m, 31.07.1954,
  • D. 23763 (ANK, E); Van: 8 km from Van to Erçek, stony slopes, 2100 m, 06.06.1966, D. 44424 (ISTO, E); Van:
  • Çatak, Kavussahap mountain, 3100 m, 23.07.1954, D. 23122 (E); Van: Gevaş, Artos mountain, rocky slopes, 2134 m, 07.1954, D. 22681 (E); Van: 36 km from Başkale to
  • Hoşap, Güzel Dere pass, rocky-steppe slopes, 2750 m, 001966, D. 45989 (E); Van: Başkale, Işıklı village, 01984, T. Baytop (ISTE); Van: Erek mountain, 001988, Özçelik & Ay (EGE); Van: Erek mountain, 3100 m, Kronenburg 130; B10 Ağrı: 3 km E of Doğubeyazıt, calcareous stones, 1750 m, 31.05.1966, D. 43945 (ISTO, E);
  • C5 Adana: Gülek, 2000 m, Sorger 62-71-23; C5 Mersin:
  • Yıldız hill, SE slopes, 2700–3000 m, 17.07.1989, Y. Gemici 4860 (EGE); C5 Niğde/İçel: Bolkar mountain, W of
  • Mededsiz hill, stony slopes, 2500–2600 m, 26.07.1984,
  • Strid et al.. 24036 (EGE); C6 Maraş: Berit mountain, 1700– 2500 m, Hausskn.; C9 Van: Harefta mountain, Başkale–
  • Koçanis, 2700–2800 m, Nábělek 3737; C10 Hakkari: Cilo mountain, 10 km W of Cilo hill, 3700 m, 09.08.1954, D. 24216 (ANK). Scorzonera sandrasica: C2 Muğla: Köyceğiz,
  • Sandras mountain, Beşparmak hill, around fire tower, steppe areas, 1970 m, 13.04.2004, Varol 5634 (GAZI);
  • Muğla: Sandras mountain, W of the summit, 1970 m, 02004, N 37 04 E 028 50, Hartvig (izo. EGE); C2
  • Muğla: Köyceğiz, Beşparmak hill, around fire tower, rocky slopes, forest boundaries, 2025 m, 23.06.2010, Makbul &
  • Coşkunçelebi 232 (RUB, KTUB). Scorzonera sublanata: A1 Çanakkale: Eceabat, Yalova village, Cistus shrubs, hill, 110 m, 02.06.2011, Makbul & Coşkunçelebi 288 (RUB, KTUB); A1(E) Çanakkale: S of Eceabat, 300 m, Sorger 63- 7-14; Çanakkale: Gökçeada, Tuzburnu–Hacıahmet, 70 m, 01976, Ö. Seçmen 1575-1588 (EGE); 60 m, 05.05.1975,
  • Ö. Seşmen 428 (EGE); A1(A) Çanakkale: Kemerdere, Sint. 1883: 325; A4 Ankara: Çubuk, Ovacık, Saraycık village,
  • Atcameydan hill, shrubby slopes, 1250–1380 m, 01992, Dündar 495 (GAZI); B1 Balıkesir: Ayvalık,
  • Cunda island, above Patriçe, 10 m, 07.05.1996, K.Alpınar
  • (ISTE); Balıkesir: Ayvalık, Armutçu–Sakarya, 20.04.1997,
  • K.Alpınar (ISTE); Balıkesir: Ayvalık, Dolap island– Mitralyöz, 5 m, 23.05.1997, K.Alpınar (ISTE); Balıkesir:
  • Ayvalık, Hakkıbey S of island, 15.04.1998, K.Alpınar
  • (ISTE); İzmir: Gümüldür, W of Değirmendere, 15.04.1969,
  • Fitz & Spitz.; İzmir: Karaburun, Balıkova–Gerence, 001980, L.Bekat 240 (EGE); İzmir: Değirmendere–
  • Gümüldür, 15.04.1969, K.Fitz (EGE); Manisa: Spil mountain, rocky and grassy, Juniperus clearance, 1224 m, 02011, Makbul & Coşkunçelebi 280 (RUB, KTUB);
  • Manisa: Spil mountain, steppe, 1196 m, 21.06.2010,
  • Makbul & Coşkunçelebi 228 (RUB, KTUB); B2 Uşak: Uşak surroundings, 900 m, Bal.; B4 Ankara: Ankara, 1100 m,
  • Balls 1933: 221; Ankara: Şereflikoçhisar–Aksaray road, 10 km from Şereflikoçhisar, steppe, 935 m, 24.05.2010, N 38 49 760 E 033 35 921, Makbul & Coşkunçelebi 199 (RUB, KTUB); C2 Aydın; below Karacasu, 400–500 m, maquis, 01965, D 41633 (E); Aydın: Aydın–Muğla, Pinus clearance, 458 m, 19.05.2011, Makbul & Coşkunçelebi 275
  • (RUB, KTUB); Muğla: Yılanlı mountain, rocky area, 1369 m, 05.2011, Makbul & Coşkunçelebi 276 (RUB, KTUB);
  • Denizli: Pamukkale, 07.05.1969, Fitz & Spitz.; Denizli:
  • Babadağ, Karacasu, above Yenice, P. brutia forest, 200 m, 01998, S.Oluk (EGE); Antalya: Elmalı, Bourgeau 169;
  • C3 Antalya: Bük forest, Kayran, Bozakman & Fitz 149; Antalya: Kumluca, W of Adrasan village, serpentine slopes, P. brutia forest, 10–50 m, 29.04.1980, Peşmen 4881
  • (HUB); Antalya: Kumluca, Adrasan village, Musa mountain, Kızıl area, serpentine slopes, P. brutia forest, 0–150 m, 23.03.1979, Peşmen 4243 (HUB); Antalya:
  • Kemer, Olimpos, 50 m, 23.03.1979, Peşmen 4229 (HUB).
  • Scorzonera tomentosa: A4 Ankara: Kalecik, 1 km W of the village, serpentine, 12.07.1998, Adıgüzel 1899 (E); Ankara:
  • Çubuk, Karagöl, Yaşmağın hill, 1500 m, 05.08.1974, Erik 683 (HUB); Çankırı: Çankırı, 800 m, Bornm. 1929: 14326;
  • A5 Amasya: Direkli village, Çetinkaya, 1550 m, 03.10.1976,
  • K.Akpınar (ISTE); Kastamonu: Tosya, Sint. 1892: 4721; A6
  • Sivas: Yıldız mountain, Cırcır, steppe, 1400 m, 14.08.1967,
  • Tobey 2312 (E); A7 Gümüşhane: Kop village, rocky slopes, 1300 m, 19.06.1967, Tobey 2043 (E); A8 Artvin: Artvin,
  • Barevan, Woronow 6153; Bayburt: Kop mountain, roadside, 2148 m, 05.07.2007, Makbul 10 (RUB, KTUB);
  • Bayburt: Bayburt–Aşkale, Kop pass, 2350 m, 16.08.1984,
  • Leblebici (EGE); Erzurum: 4 km W of Kuzgun village, rocky slopes, 2250–2350 m, 19.07.1977, Tatlı 5857 (HUB);
  • Gümüşhane: near Bayburt, Bourgeau 147; A9 Kars: E of Kağızman mountain, 17 km from Akçay to Cumaçay, 2070 m, 07.1966, D. 46720 (ISTO, E); Kars: Demirkapı,
  • Woronow 12686; B2 Uşak: Bulkas mountain, 1250 m, Bal.;
  • B3 Konya: Sultan mountain, 28.06.1953, Birand & M.
  • Zohary; B4 Ankara: near Ankara, Liston; Ankara: Mamak, Kıbrıs village valley, stony slopes, 1000–1100 m, 01.07.2004,
  • Aslan 1592 (GAZI); Ankara: Beynam, 400 m, 05.09.1949,
  • D. 13101; B5 Yozgat: Yıldızlı–Akdağmadeni, 10 km to
  • Akdağmadeni, stony area, 1113 m, 06.06.2010, Makbul &
  • Coşkunçelebi 211 (RUB, KTUB); B6 Adana: Binboğa mountain, Yalak, 2000 m, 14.07.1952, D. 19938 (ANK);
  • Kayseri: Sarız, Yalak, Binboğa mountain, above Yalak, rocky field, 1700–2000 m, 08.07.1992, Aytaç & Duman 5242 (GAZI); Kayseri: Akkışla, above Ganişeyk, Hınzır mountain, 1900 m, 20.08.1979, Çelik 771 (HUB); Kayseri,
  • Sarız, Yalak, Binboğa mountain, stream side, stony field, 1500–1700 m, 04.08.1991, Aytaç 4349 (GAZI); Kayseri:
  • Pınarbaşı, Kaynar, S slopes of Hınzır mountain, above Kara Ahmed, slopes, 2000 m, 18.07.1975, Çelik 531
  • (ANK); Kayseri: Sarız, Yeşilkent, Binboğa mountain, climbing to Tekke stone, calcareous stones, 1734 m, 02010, Makbul & Coşkunçelebi 249 (RUB, KTUB);
  • Malatya: Doğanşehir, Eskiköy, Çobanderesi-İhtiyarcave, Juniperus excelsa–Quercus forest, 1600–1800 m, 01971, Peşmen 2635 (HUB); Malatya: Doğanşehir,
  • Dedeyazı village, Keşiş mountain, alpine steppe, 1800– 2200 m, 28.07.1971, Peşmen 2580, (HUB); Malatya:
  • Akçadağ, Kürecik, Aksüt village, 1443 m, 18.07.2005, Y.
  • Yeşil (ISTE); Maraş: Göksun, Kınıkköz village, Gözpınarı mountain, 1700 m, 22.08.1977, Yıldız 1532 (HUB); Maraş:
  • Göksun–Değirmendere, Koruyaz mountain–Meryemçil protected area, 1750–2000 m, 05.07.1993, Ekici 1577
  • (GAZI); Maraş: Göksun, Binboğa mountain pass, 1500 m, 01952 (E); Sivas: Ak mountain, near Sivas, 2200 m,
  • Siehe 1911: 308; Sivas: Şarkışla, 1500 m, D. 32720 (E);
  • Sivas: Hekimhan–Kangal, hillside, 1800 m, 02.09.1954, D. 24844 (E); Sivas: 8 km from Sivas to Ulaş, 1300–1450 m, 001982, D. 68628 (HUB); Sivas: Hafik, Koşutdere–
  • Bakışecik, stony places, 1250 m, 30.06.1991, Aydoğdu 3245 (GAZI); Sivas: Şarkışla, Evcidere Kale–Alaman, calcareous stones, 1400–1500 m, 19.07.1979, Ekim &
  • Düzenli 4022 (ANK); Sivas: Ulaş, Tecer village, Tecer mountain, 1600 m, 21.07.1978, A.Baytop et al. (ISTE);
  • Sivas: Çelebiler, Okatan farmer sulfated soils, 1500 m, 001972, Y.Okatan (ISTF); Sivas/Tokat: Çamlıbel pass, 1650 m, 21.07.1978, A.Baytop et al. (ISTE); B7 Elazığ: Kop mountain, Sarık village, steppe, 1700 m, 18.07.1984, Evren 2108 (GAZI); Erzincan: Kemaliye (Eğin), Sırakonaklar village, calcareous stones, 1600 m, 10.07.1982, Tanker 10516 (ANK); Erzincan: Sipikör, Sint. 1890: 3309;
  • Erzincan: Keşiş mountain, volcanic slopes, 2600 m, 01957, D. 31759 (E); Erzincan: Erzincan–Refahiye, 25 km E of Refahiye, near Sakallutan pass, serpentine area, 1960–2020 m, 15.08.2001, N 39 52 146 E 039 04 054,
  • Adıgüzel 4156 (GAZI, E); Erzurum: W of Erzurum,
  • Erzurum–Kandilli, Ağaver–Kandilli, Tapi stream, limy slopes, 1960 m, 17.07.1969, O.Özbay (ISTF); Malatya:
  • Kırlangıç village, Eğri Dere, back of the cave, 1500 m, 01993, N.Gören (ISTE); Malatya: Doğanşehir,
  • Eskiköy, above İhtiyar cave, alpine steppe, 1900–2000 m, 01971, Peşmen 2331 (EGE); Tunceli: Ovacık, Munzur mountain, Karagöl valley, 1500 m, 09.09.1979, Yıldırımlı 2463 (HUB); Tunceli: Munzur mountain, above Ovacık,
  • Aksu stream, stony places, 1800 m, 21.07.1957, D. 31470
  • (E); 2100 m, 19.07.1957, D. 31313 (E); Tunceli: N pass of
  • Pülümür, near Küçü Dere, rocky slopes, 03.08.1974, Rix 2388 (E); Tunceli/Erzincan: Pülümür pass, grassland, 1950 m, 07.197, F. Holtz (EGE); B8 Erzurum: Palandöken mountain, Kırkdeğirmenler, 1950–2100 m, 22.07.1984,
  • Demirkuş 2028 (HUB); Erzurum: Tercan–Selepur, 1400 m, 07.1957, D. 30954 (E); Erzurum: 66 km from Hınıs to Erzurum, Aras pass, 1650 m, 12.07.1966, D. 46437
  • (ISTO, E); C3 Antalya: Gazipaşa–Çayıryaka plateau road, rocky-stony places, 1714 m, 25.07.2010, Makbul &
  • Coşkunçelebi 254 (RUB, KTUB); Konya: Beyşehir, Kurucuova, Radar-Karagöl, calcareous stones, alpine, 2000–2500 m, 24.07.1975, Güner 2290 (HUB); Konya:
  • Kuru tepe, 1400 m, Sorger 66-44-105; C4 İçel: Anamur,
  • Anamur–Kazancı road, Kızıl Alan, 1300 m, 24.06.1984,
  • Sümbül 3103 (HUB); Konya: 19 km from Konya to
  • Beyşehir, 1210 m, Hub.-Mor. 8694; C5 Konya: Ereğli,
  • Aydos mountain, Berendi, Boğaz, calcareous stones, 1600 m, 06.1976, Erik 1705 (HUB); C6 Osmaniye: Zorkun,
  • Amanos mountain, Keldaz Hill, calcareous stones, 2020 m, 002010, Makbul & Coşkunçelebi 241 (RUB, KTUB);
  • C7 Malatya: 74 km SW of Malatya, 1550 m, Sorger 71-45
  • Scorzonera tuzgoluensis: B4 Konya: Cihanbeyli, Gölyazı–Tuzgölü, 908 m, 11.06.2009, N 38 32 552 E 033 21 188, A. Duran 8368 (holo.: KNYA, iso.: GAZI, ANK, HUB, MR); 07.2006, A. Duran 7241 (KNYA, MR); 29.06.2009,
  • B. Doğan 2100 (KNYA, GAZI, MR); Konya: Cihanbeyli, climbing to Gölyazı plateau, Dumanağlı plateau, salty swamp, 908 m, 06.07.2011, Makbul & Coşkunçelebi 313
  • (RUB, KTUB). Scorzonera ulrichii: C4 Antalya: Alanya, Çökele–Gökbel, calcareous area, near Pinus forest, 1425 m, 007.2003, Göktürk 5107 (HUB, GAZI); Antalya:
  • Alanya, Mahmutlar–Hadim road, c. 35 km NE of Mahmutlar and 15 km S Çayarası, 1 km N Elmalısu, gravely, rocky slopes with Pinus nigra var. caraminaca forest clearance, W-exp., limestone, 1250 m, 11.06.2002,
  • Robert Ulrich (Tubingen) 2/12 (holo.:B, iso.: E, ISTE); C4 Antalya: Alanya, Mahmutlar, Elmalısu–Hadim road, after Kuşkaya, calcareous rocks, under Pinus nigra, 1336 m, 002010, Makbul & Coşkunçelebi 237 (RUB, KTUB);
  • Antalya: Alanya, Mahmutlar–Hadim road, Elmalısu, junction of Gökbel plateau road, calcareous rocks, under Pinus nigra, 1336 m, 05.07.2010, Makbul & Coşkunçelebi 238 (RUB, KTUB). Scorzonera veratrifolia: B9 Bitlis:
  • Kambos mountain, above Hurmuz, 1830 m, 31.06.1954, D. 23494 (E) ; Van: Pelli mountain, 54 km SW of Van 2500 m, 001971, Edmondson 618 (E) ; Van: Van lake, 20 km SE of Tatvan, 1960 m, Frodin 1939, 1971 and 1972; C9
  • Hakkari: between Beytuşşebap and Uludere, W of Tanin mountain, 2500 m, 25.07.1974, Rix 2338 (E) ; Hakkari:
  • Cilo mountain, Diz stream, 1740 m, 06.08.1954, D. 23926
  • (E) ; Van: Çatak, Dalbastı village, 1350–1600 m, 20.07.2002,
  • Bani 1259 (GAZI) ; C10 Hakkari: Sat mountain, 2000– 2100 m, 30.06.1966, D. 45812 (ISTO, E) ; Hakkari Cilo mountain, 2500 m, 25.08.2011, MF 27719 ; Hakkari:
  • Koçanis, 2500 m, 21.07.2011, MF 27566 ; Hakkari:
  • Esendere, Mor mountain, 2700 m, 24.07.2011, MF 27194 ;
  • Hakkari: Diz stream, 2200 m, 23.07.2011, MF 27168 .
  • Scorzonera yildirimlii: C6 Osmaniye: Amanos mountain, Zorkun plateau, Keldaz hill, 1950 m, 05.07.2001, N 36 58 95 E 036 24 22, Duran 5765-6824 (iso.: HUB, GAZI; para.: E); Osmaniye: Zorkun, Amanos mountain, Keldaz hill,
  • Halep Gösteren, calcareous rocky hills, 2073 m, 05.07.2010,
  • Makbul & Coşkunçelebi 243 (RUB, KTUB).