Oligocene vegetation and climate characteristics in north-west Turkey: data from the south-western part of the Thrace Basin

In this paper we present the first palynomorph and mollusc assemblages from the sediments in three different sections. From east to west, these are the Şevketiye (northern Biga Peninsula), the Tayfur (Gelibolu Peninsula) and the Kuzu harbour (Gökçeada) (parts of the Danişmen Formation) sections in the south and south-western side of the Thrace Basin (north-west Turkey), with the aim of obtaining information about the composition and structure of vegetation and climate during the Oligocene. The stratigraphic interval extends from late Rupelian to Chattian. The Danişmen Formation in the Şevketiye section yielded a palynomorph association with abundant coastal palms (Arecaceae; Lepidocaryoidae), and mangrove pollen (Pelliciera). A similar assemblage from the Kuzu harbour section was also obtained, with minor contributions of mangrove elements Nypa and Acrostichum aureum, Arecaceae type palm, undifferentiated dinoflagellate cysts and microforaminiferal linings. These palynomorph assemblages, combined with the mollusc data, indicate that low-lying coastal environments prevailed. In contrast, the palynomorphs from the Tayfur section represent a non-marine environment lacking mangrove elements, palm trees, dinoflagellate cysts and microforaminiferal linings. The diversity of angiosperm taxa in the Tayfur palynoflora, which form the bulk of the assemblage, indicates terrestrial vegetation. Quantitative palaeoclimate analyses are based on the Coexistence Approach method, and yield over 22 °C at the coast as indicated by mangrove elements and palms in the Şevketiye and Kuzu harbour palynofloras. For the Tayfur palynoflora, mean annual temperature ranged between 16.5 and 21.3 °C. This indicates a climate cooling, corresponding to the transition from Rupelian to Chattian, and resulted in the pollen changes from mangrove bearing coastal deposits to more inland vegetation.

Oligocene vegetation and climate characteristics in north-west Turkey: data from the south-western part of the Thrace Basin

In this paper we present the first palynomorph and mollusc assemblages from the sediments in three different sections. From east to west, these are the Şevketiye (northern Biga Peninsula), the Tayfur (Gelibolu Peninsula) and the Kuzu harbour (Gökçeada) (parts of the Danişmen Formation) sections in the south and south-western side of the Thrace Basin (north-west Turkey), with the aim of obtaining information about the composition and structure of vegetation and climate during the Oligocene. The stratigraphic interval extends from late Rupelian to Chattian. The Danişmen Formation in the Şevketiye section yielded a palynomorph association with abundant coastal palms (Arecaceae; Lepidocaryoidae), and mangrove pollen (Pelliciera). A similar assemblage from the Kuzu harbour section was also obtained, with minor contributions of mangrove elements Nypa and Acrostichum aureum, Arecaceae type palm, undifferentiated dinoflagellate cysts and microforaminiferal linings. These palynomorph assemblages, combined with the mollusc data, indicate that low-lying coastal environments prevailed. In contrast, the palynomorphs from the Tayfur section represent a non-marine environment lacking mangrove elements, palm trees, dinoflagellate cysts and microforaminiferal linings. The diversity of angiosperm taxa in the Tayfur palynoflora, which form the bulk of the assemblage, indicates terrestrial vegetation. Quantitative palaeoclimate analyses are based on the Coexistence Approach method, and yield over 22 °C at the coast as indicated by mangrove elements and palms in the Şevketiye and Kuzu harbour palynofloras. For the Tayfur palynoflora, mean annual temperature ranged between 16.5 and 21.3 °C. This indicates a climate cooling, corresponding to the transition from Rupelian to Chattian, and resulted in the pollen changes from mangrove bearing coastal deposits to more inland vegetation.

___

  • Abreu, V.S. & Anderson, J.B. 1998. Glacial eustasy during the Cenozoic: sequence stratigraphic implications. American Association of Petroleum Geologists Bulletin 82, 1385–1400.
  • Akgün, F. 2002. Stratigraphic and paleoenvironmental significance of Eocene palynomorphs of the Çorum–Amasya area in the central Anatolia, Turkey. Acta Palaeontologica Sinica 41, 576– 5
  • Akgün, F., Akay, E. & Erdoğan, B. 2002. Terrestrial to Shallow Marine Deposition in Central Anatolia: A Palynological Approach. Turkish Journal of Earth Sciences 11, 1–27.
  • Akkiraz, M.S. 2000. Palynostratigraphy of the Oligo-Miocene Denizli
  • Çardak Molasse Basin. Master Thesis, Dokuz Eylül Üniversitesi, İzmir–Türkiye [unpublished]. Akkiraz, M.S. & Akgün, F. 2005. Palynology and age of the Early Oligocene units in Çardak–Tokça Basin, Southwest Anatolia: Paleoecological implications. Geobios 38, 283–299.
  • Akkiraz, M.S., Akgün, F., Örçen, S., Bruch, A.A. & Mosbrugger, V. 200 Stratigraphic and Palaeoenvironmental Significance of Bartonian–Priabonian (Middle–Late Eocene) Microfossils from the Başçeşme Formation, Denizli Province, Western Anatolia. Turkish Journal of Earth Science 15, 155–180. Akkiraz, M.S.,  Kayseri, M.S. &  Akgün, F. 2008.  Palaecology of Coal-Bearing Eocene Sediments in Central Anatolia (Turkey) Based on Quantitative Palynological Data. Turkish Journal of Earth Sciences 17, 317–360.
  • Akkiraz, M.S., Akgün, F. & Örçen, S. 2011. Stratigraphy and palaeoenvironment of the Lower–“middle” Oligocene units in the northern part of the Western Taurides (İncesu area, Isparta, Turkey). Journal of Asian Earth Sciences 40, 452–474.
  • Akyol, E, 1971. Microflore de Oligocène inférieur récoltée dans un sondage près d’Avcikoru, Şile–İstanbul: Pollen et Spores 13, 117–133.
  • Bande, M.B. & Prakash, U. 1986. The Tertiary flora of Southeast Asia with remarks on its paleoenvironmental and phytogeography of the Indo-Malayan region. Review of Palaeobotany and Palynology 49, 203–233.
  • Batı, Z. 1996. Palynostratigraphy and Coal Petrography of the Upper Oligocene Lignites of the Northern Thrace Basin, NW Turkey. PhD Thesis, Middle East Technical University, Ankara–Turkey [unpublished].
  • Bozukov, V., Utescher, T. & Ivanov, M. 2009. Late Eocene to early Miocene climate and vegetation of Bulgaria. Review of Palaeobotany and Palynology 153, 360–374.
  • Cavagnetto, C. & Anadón, P. 1995. Une mangrove complexe dans le Bartonien du basin de le l’Ebre (NE de l’Espagne). Palaeontographica Abteilung B Ionnides 236, 147–165.
  • Cavagnetto, C. & Anadón, P. 1996. Preliminary palynological data on floristic and climatic changes during the Middle Eocene–Early Oligocene of the eastern Ebro Basin, northeast Spain. Review of Palaeobotany and Palynology 92, 281–305.
  • Collinson, M.E. & Hooker, J.J. 2003. Palaeogene vegetation of Eurasia: framework for mammalian faunas. Deinsea 10, 41–83.
  • Coşkun, B. 2000. North Anatolian Fault–Saroz Gulf relationship and their relevance to hydrocarbon exploration, North Aegean Sea Turkey. Marine and Petroleum Geology 17, 751–772.
  • Ediger, V.Ş., Batı, Z. & Alişan, C. 1990. Paleopalynology and Paleoecology of Calamus like Disulcate Pollen Grains. Review of Paleobotany and Palynology 62, 97–105.
  • El Beıaly, S.Y. 1998. Stratigraphic and palaenvironmental significance of Eocene palynomorphs from the Rusayl Shale Formation, Al Khawd, northern Oman. Review of Palaeobotany and Palynology 102, 249–258.
  • Elsik, W.C., Ediger, V.Ş. & Batı, Z. 1990. Fossil Fungal spore: Anatolinites gen. nov. Palynology 14, 91–103.
  • Erginal, A.E., Kıyak, N.G., Bozcu, M., Ertek, A., Güngüneş, H., Sungur, A. & Türker, G. 2008. On the Origin and Age of the Arıburnu Beachrock, Gelibolu Peninsula, Turkey. Turkish Journal of Earth Sciences 17, 803–819.
  • Fechner, G.G. 1988. Selected Palynomorphs from the Lower to Middle Eocene of the South Atlas Border Zone (Morocco) and their Environmental Significance. Palaeogeography, Palaeoclimatology, Palaeoecology 65, 73–79.
  • Frederiksen, N.O. 1980. Sporomorphs from the Jackson Group (Upper Eocene) and adjacent strata of Mississipi and western Alabama. Geological Survey Professional Paper 1084, 1–75.
  • Frederiksen, N.O. 1985. Review of early Tertiary sporomorph palaeoecology. American Association of Stratigraphical Palynologists Contribution Series 19, 1–92.
  • Frederiksen, N.O. 1988. Sporomorph biostratigraphy, floral changes, and paleoclimatology, Eocene and earliest Oligocene of the eastern Gulf Coast. United States Geological Survey Professional Paper 1448, 1–68.
  • Frederiksen, N.O. 1994. Paleocene floral diversities and turnover events in eastern North America and their relation to diversity models. Review of Palaeobotany and Palynology 32, 225–238.
  • Germeraad, J.H., Hopping, C.A. & Müller, J. 1968. Palynology of Tertiary sediments from tropical areas. Review of Palaeobotany and Palynology 6, 189–348.
  • Görür, N. & Okay, A.İ. 1996. Fore-arc origin of the Thrace Basin, northwest Turkey. Geologische Rundschau 85, 662–668.
  • Graham, A. 1977. New records of Pelliciera (Theaceae/Pelliceriaceae) in the Tertiary of the Caribbean. Biotropica 9, 48–52.
  • Graham, A. 1995. Diversification of Gula/Caribbean Mangrove Communities through Cenozoic time. Biotropica 27, 20–27.
  • Gruas-Cavagnetto, C. 1977. Etude palynologique de l’Eocene du Bassin Anglo-Parisien. These de Doctorate d’Etat es Sciences Naturelles, Universite de Pierre Marie Curie VI, 287p.
  • Gürgey, K., Philp, R.P., Clayton, C., Emiroğlu, H. & Siyako, M. 200 Geochemical and isotopic approach to maturity/source/ mixing estimations for natural gas and associated condensates in the Thrace Basin, NW Turkey. Applied Geochemistry 20, 2017–2037.
  • Haq, B., Hardenbol, J. & Vail, P. 1987. The chronology of fluctuating sea levels since the Triassic. Science 235, 1156–1166.
  • Hornibrook, N.B. 1992. New Zealand Cenozoic marine paleoclimates: a review based on the distribution of some shallow water and terrestrial biota. In: Tsuchi, R. & Ingles, J. (eds), Pacific Neogene
  • Environment, Evolution and Events. University of Tokyo Press 83–106. Hoşgörmez, H. & Yalçın, M.N. 2005. Gas-source rock correlation in Thrace basin, Turkey. Marine and Petroleum Geology 22, 901–916.
  • Hoşgörmez, H., Yalçın, M.N., Cramer, B., Gerling, P. & Mann, U. 2005.
  • Molecular and isotopic composition of gas occurrences in the Thrace basin (Turkey): origin of the gases and characteristics of possible source rocks. Chemical Geology 214, 179–191. Huvaz, O., Karahanoğlu, N. & Ediger, V. 2007. The Thermal Gradient
  • History of the Thrace Basin, NW Turkey: Correlation with Basin Evolution Processes. Journal of Petroleum Geology 30, 3–24 Ivanov, D., Ashraf, A.R., Mosbrugger, V. & Palamarev, E. 2002. Palynological evidence for Miocene climate change in the Forecarpathian Basin (Central Paratethys, NW Bulgaria). Paleogeography Paleoclimatology Paleoecology 178, 19–37.
  • Ivanov, D., Ashraf, A.R. & Mosbrugger, V. 2007. Late Oligocene and Miocene climate and vegetation in the Eastern Paratethys area
  • (northeast Bulgaria), based on pollen data. Paleogeography Paleoclimatology Paleoecology 255, 34–360. İslamoğlu, Y., Harzhauser, M., Gross, M., Jiménez-Moreno, G., Coric, S., Kroh, A., Rögl, F. & Van Der Made, J. 2010. From Tethys to Eastern Paratethys: Oligocene depositional environments, paleoecology and paleobiogeography of the Thrace Basin (NW Turkey). International Journal of Earth Sciences 99, 183–200.
  • Jiménez-Moreno, G., Mandic, O., Harzhauser, M., Pavelić, D. & Vranjković, A. 2008. Vegetation and climate dynamics during the early Middle Miocene from Lake Sinj (Dinaride Lake System, SE Croatia). Review of Palaeobotany and Palynology 152, 237–245.
  • Jiménez-Moreno, G., Rodriguez-Tovar, F.J., Pardo-Iguzquiza, E., Fauquette, S., Suc, J.-P. & Müller, P. 2005. High-resolution palynological analysis in late early–middle Miocene core from the Pannonian Basin, Hungary: climatic changes, astronomical forcing and eustatic fluctuations in the Central Paratethys. Palaeogeogaphy, Palaeoclimatology, Palaeoecology 216, 73–97.
  • Kantarcı, M.D. 2011. Vertical climate zones in Biga peninsula: The impact of climate change and air pollution on forests. Procedia Social and Behavioral Sciences 19, 797–810.
  • Kavgacı, A., Čarni, A., Tecimen, B. & Özalp, G. 2010. Diversity and Ecological Differentiation of Oak Forests in NW Thrace (Turkey). Archives of Biological Sciences Belgrade 62, 705–718.
  • Kayseri, M.S. 2009. Milas–Ören Havzasının Geç Rüpeliyen-Erken Şattiyen Dönemindeki Paleoiklim ve Palaeovejetasyonu [Palaeoclimate and Palaeovegetation in the Milas–Ören Basin During the Late Rupelian-Early Chattian Period]. Abstracts, Türkiye jeoloji Kurultayı, Ankara, p. 742–743 [in Turkish and English].
  • Kesgin, Y. & Varol, B. 2003. Gökçeada ve Bozcaada’nın Tersiyer Jeolojisi (Çanakkale), Türkiye, Maden Tetkik ve Arama Dergisi 126, 49–67 [in Turkish].
  • Kopp, K.O., Pavoni, N. & Schindler, C. 1969. Geologie Thrakiens IV: Das Ergene-Becken. Beihhefte zum Geologischen Jahrbuch Heft 76, 1–71.
  • Less, G.Y., Özcan, E. & Okay, A.İ. 2011. Stratigraphy and larger Foraminifera of the Middle Eocene to Lower Oligocene shallow-marine units in the northern and eastern parts of the Thrace Basin, NW Turkey. Turkish Journal of Earth Sciences 20, 793–845.
  • Morton, B. 1983. Mangrove Bivalves. In: Wilbur, K.W (ed), The Mollusca. Academic Press, Orlando 6, 77–138.
  • Mosbrugger, V. & Utescher, T. 1997. The coexistence approach–a method for quantitative reconstructions of Tertiary terrestrial palaeoclimate data using the plant fossils. Palaeogeography, Palaeoclimatology, Palaeoecology 134, 61–86.
  • Mosbrugger, V., Utescher, T. & Dilcher, D.L. 2005. Cenozoic continental climatic evolution of Central Europe. Proceedings of the National Academy of Sciences of the United States of America (PNAS) 102, 14964–14969.
  • Mudie, P.J., Rochon, A. & Aksu, A.E. 2002. Pollen stratigraphy of Late Quaternary cores from Marmara Sea: land–sea correlation and paleoclimatic history. Marine Geology 190, 233–260.
  • Müller, J. 1980. Fossil pollen records of extant angiosperms. Botanical Review, 47, 1–142.
  • Müller, J. 1981. Palynological evidence for Paleogene climatic changes. Mémoires du Museum National d’Historie Naturelle, Nouvelle Séries, B Botanique XXVII, 211–218.
  • Nakoman, E. 1968. Contribution a l’étude de la microflore Tertiaire des lignites de Seyitömer (Turquie). Pollen et Spores 10, 521– 5
  • Nickel, B. 1996. Palynofazies und Palynostratigraphie der Pechelbronn Schichten im nördlichen Oberrheintalgraben. Palaeontographica Abteilung B Ionnides 240, 1–151.
  • Okay, A.İ., Özcan, E., Cavazza, W., Okay, N. & Less, G.Y. 2010. Basement types, Lower Eocene Series, Upper Eocene Olistostromes and the Initiation of the Southern Thrace Basin, NW Turkey. Turkish Journal of Earth Sciences 19, 1–25.
  • Özcan, E., Less, G.Y., Okay, A.İ., Baldi-Beke M., Kollanyi K. & Yılmaz, İ.Ö. 2010. Stratigraphy and larger foraminifera of the Eocene shallow-marine and olistostromal units of the southern part of the Thrace Basin, NW Turkey. Turkish Journal of Earth Sciences 19, 27–77.
  • Palamarev, E., Kitanov, G., Staneva, K. & Bozukov, V. 2000. Fossil flora from Paleogene sediments in the northern area of the Mesta Graben in the Western Rhodopes. II. Analysis and stratigraphic importance of the flora. Phytologia Balcanica 6, 3–
  • Peel, M.C., Finlayson, B.L. & McMahon, T.A. 2007. Updated world map of the Köppen-Geiger climate classification. Hydrology and Earth System Sciences 11, 1633–1644.
  • Plaziat, J.C., Cavagnetto, C., Koeniguer, J.C. & Baltzer, F. 2001. History and biogeography of the mangrove ecosystem, based on a critical reassessment of the paleontological record. Wetlands Ecology and Management 9, 161–179.
  • Pole, M.S. & Macphail, M.K. 1996. Eocene Nypa from Regatta Point, Tasmania. Review of Palaeobotany and Palynology 92, 55–67.
  • Riegel, W., Bode, T., Hammer, J., Hammer-Schiemann, G., Lenz, O. & Wilde, V. 1999. The palaeoecology of the Lower and Middle
  • Eocene at Helmstedt, northern Germany – a study in contrasts. Acta Palaeobotanica Supplement 2, 349–358. Roberts, N. & Wright, JR.H.E. 1993. Vegetation, Lake–Level, and Climatic History of the Near East and Southwest Asia. In: Wright, Jr. H.E. Kutzbach, J. E. Web III. T. Ruddiman, W.F. Street-Perrot, F.A. & Bartlein, P.J. (eds), Global Climates Since the Last Glacial maximum. University of Minnesota Press, USA 194–220.
  • Rögl, F. 1998. Palaeogeographic considerations for Mediterranean and Paratethys Seaways (Oligocene to Miocene). Annalen des Naturhistorischen Museums in Wien 99, 279–310.
  • Rögl, F. 1999. Short Note Mediterranean and Paratethys. Facts and Hypotheses of An Oligocene To Miocene Palaeogeography (Short Overview). Geologica Carpathica 50, 339–349.
  • Rull, V. 1998. Middle Eocene mangroves and vegetation changes in the Maracibo Basin. Palaios 13, 287–296.
  • Rull, V. 1999. Paleofloristic and paleovegetational changes across the Paleocene-Eocene boundary in northern South America. Palaeogeography, Palaeoclimatology, Palaeoecology 107, 83–95.
  • Sakınç, M., Yaltırak, C. & Oktay, F.Y. 1999. Palaeogeographic evolution of the Thrace Neogene Basin and the TethysParatethys relations at northwestern Turkey (Thrace). Palaeogeography Palaeoclimatology Palaeoecology 153, 17–40.
  • Sırdaş, S. & Şen, Z. 2003. Spatio-temporal drought analysis in the Trakya region, Turkey, Hydrological Sciences Journal 48, 809– 8
  • Siyako, M. 2003. Trakya bölgesinin Tersiyer litostratigrafi birimleri. In: Okay, A.İ., Siyako, M. & Yurtsever, A. (eds.), Trakya bölgesinin Litostratigrafisi. Mineral Research and Exploration Institute of Turkey (MTA), Ankara.
  • Siyako, M. & Huvaz, O. 2007. Eocene stratigraphic evolution of the Thrace Basin, Turkey. Sedimentary Geology 198, 75–91.
  • Suc, J.-P. 1984. Origin and Evolution of the Mediterranean Vegetation and Climate in Europe. Nature 307, 429–432.
  • Srivastava, S.K. & Binda, P.L. 1991. Depositional History of the Early Eocene Shumaysi Formation, Saudi Arabia. Palynology 15, 47–
  • Thanikaimoni, G. 1987. Mangrove Palynology. Travaux de la Section Scientifique et Technique. French Institute of Pondichéry 24, 1–
  • Thanikaimoni, G., Caratini, C., Venkatachala, B.S., Ramanujam, C.G.K. & Kar, R.K. 1984. Selected Tertiary angiosperm pollens from India and their relationship with African Tertiary pollen, Travaux de la Section Scientifique et Technique XIX, 1–192.
  • Turgut, S., Türkaslan, M. & Perinçek, D. 1991. Evolution of the Thrace sedimentary basin and its hydrocarbon prospectivity. In: Spencer, A.M. (ed), Generation, Accumulation, and Production of Europe’s Hydrocarbons. Special Publication of European Association of Petroleum Geoscientists 1, 415–437.
  • Turgut, S. & Eseller, G. 2000. Sequence stratigraphy, tectonics and depositional history in eastern Thrace Basin, NW Turkey. Marine and Petroleum Geology 17, 61–100.
  • Utescher, T., Djordjevic-Milutinovic, D., Bruch, A. & Mosbrugger, V. 200 Palaeoclimate and vegetation change in Serbia during the last 30 Ma. Palaeogeography, Palaeoclimatology, Palaeoecology 253, 157–168. Utescher, T. & Mosbrugger, V. 2007. Eocene vegetation patterns reconstructed from plant diversity – a global perspective.
  • Palaeogeography, Palaeoclimatology, Palaeoecology 247, 243– 2 Van Der Hammen, T. & Wijmstra, T.A. 1964. A palynological study on the Tertiary and Upper Cretaceous of British Guiana. Leidse
  • Geologische Mededelingen 30, 183–241. Westgate, J.W. & Gee, C.T. 1990. Palaeoecology of a Middle Eocene mangrove biota (vertebrates, plants and invertebrates) from southwest Texas. Palaeogeography, Palaeoclimatology, Palaeoecology 78, 163–177.
  • Yarcı, C. 2000. Işıklar Dağı’nın (Tekirdağ) vejetasyonu üzerinde fitososyolojik ve ekolojik araştırmalar. Erciyes Üniversitesi Fen Bilimleri Enstitüsü Dergisi 16 (1–2): 1–10 [in Turkish with English abstract].
Turkish Journal of Earth Sciences-Cover
  • ISSN: 1300-0985
  • Yayın Aralığı: Yılda 6 Sayı
  • Yayıncı: TÜBİTAK
Sayıdaki Diğer Makaleler

AFC-Modeler: a Microsoft® Excel© workbook program for modelling assimilation combined with fractional crystallization (AFC) process in magmatic systems by using equations of DePaolo (1981)

Mehmet KESKİN

Prediction of weathering development in metarhyolites of the Ilgın (Konya) area, SW Turkey

Erkan BOZKURTOĞLU, Şenel ÖZDAMAR, Hatice Ünal ERCAN

Comment on Al-in-Hornblende Thermobarometry and Sr-Nd-O-Pb Isotopic Compositions of the Early Miocene Alaçam Granite in NW Anatolia (Turkey)

Fuat ERKÜL, Sibel Tatar ERKÜL

Microfacies correlation analysis of the Oligocene-Miocene Asmari Formation, in the central part of the Rag-e-Safid anticlinal oil field, Zagros Basin, south-west Iran

Mahnaz AMIRSHAHKARAMI

Revised Late Campanian-Danian age of the melange-related turbiditic sequence in the Mersin area (Central Taurides, S. Turkey)

Hayati KOÇ, Kemal TASLI, Erol ÖZER

Comment on "Al-in-Hornblende Thermobarometry and Sr-Nd-O-Pb Isotopic Compositions of the Early Miocene Alaçam Granite in NW Anatolia (Turkey)"

Ferdi TATAR ERKÜL, Fuat ERKÜL

Tectonosedimentary development and palaeoenvironmental changes in the Acıgöl shallow-perennial playa-lake basin, SW Anatolia, Turkey

Cahit HELVACI, M. Cihat ALÇİÇEK, İbrahim GÜNDOĞAN, Ünsal GEMİCİ

Coeval Shoshonitic-ultrapotassic dyke emplacements within the Kestanbol Pluton, Ezine – Biga Peninsula (NW Anatolia)

Cüneyt AKAL

The efficacy of travertine as a palaeoenvironmental indicator: palaeomagnetic study of neotectonic examples from Denizli, Turkey

Bekir Levent MESCİ, Orhan TATAR, John D. A. PIPER, Halil GÜRSOY, Erhan ALTUNEL, Stephen CROWLEY

Tragulidae (Artiodactyla, Ruminantia) from the Middle Miocene Chinji Formation of Pakistan

Muhammad Akbar KHAN, Muhammad AKHTAR